Italian Italian mathematician and physicist, known for his contributions to mathematical biology Vito Volterra of Sapienza – Università di Roma published "Varizioni e fluttuazioni del numero d'individui in specie animali conviventi" in Mem. R. Acad. Naz. dei Lincei (ser.6) II, 31-113.
This work was translated into English and published in the journal Nature the same year as "Fluctuations in the abundance of a species considered mathematically". In this paper Volterra created the basic equations for two species interactions.