A: London, England, United Kingdom
In 1934 British crystallographer, polymath and writer John Desmond Bernal at the University of London, and British chemist and crystallographer Dorothy Crowfoot (Hodgkin), took the first X-ray photograph of a protein structure—crystalline pepsin. They showed that crystals of pepsin give an X-ray diffaction pattern, beginning protein crystallography. This may also be the beginning of structural molecular biology.
Bernal & Crowfoot, "X-Ray Photographs of Crystalline Pepsin," Nature 133 (1934) 794-95.